Benthophilinae

Benthophilinae
	The pale monkey goby (Neogobius fluviatilis) and the dark bighead goby (Ponticola kessleri) are representatives of Benthophilinae
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Gobiiformes
Family:	Gobiidae
Subfamily:	Benthophilinae; Beling & Iljin, 1927
Type genus
	Benthophilus; Eichwald, 1831
Genera
	See text

The Benthophilinae are a subfamily of gobies endemic to the Ponto-Caspian region (including the Marmara, Black, Azov, Caspian, and Aral Seas).^[1] The subfamily includes about 50 species. The representatives of the subfamily have fused pelvic fins and elongated dorsal and anal fins.^[2] They are distinguished from the closely related subfamily Gobiinae by the absence of a swimbladder in adults and location of the uppermost rays of the pectoral fins within the fin membrane.^[3]

The Catalog of Fishes still considers these fishes as belonging to the subfamily Gobiinae.^[4]

Systematics of the subfamily

Tribe Benthophilini
- Anatirostrum
- Benthophiloides
- Benthophilus (type genus)
- Caspiosoma
Tribe Neogobiini
- Neogobius (type genus)
Tribe Ponticolini
- Babka
- Mesogobius
- Ponticola (type genus)
- Proterorhinus^[5]

References

^ Simonović P.D.; Nikolić V.P.; Skóra K.E. (1996). "Vertebral number in Ponto-Caspian gobies: phylogenetic relevance". J. Fish Biol. 49 (5): 1027–1029. Bibcode:1996JFBio..49.1027S. doi:10.1111/j.1095-8649.1996.tb00098.x.
^ Miller P.J. (1986). "Gobiidae". In Whitehead P.J.P.; Bauchot M.-L.; Hureau J.-C.; Nielsen J.; Tortonese E. (eds.). Fishes of the North-eastern Atlantic and the Mediterranean Vol. 3. Paris: UNESCO.
^ Pinchuk V.I. (1991). "K voprosu o grupirovkakh vidov v predelakh roda Neogobius (Perciformes)" [On the issue of species groupings within the genus Neogobius (Perciformes)]. Voprosy Ikhtiologii (in Russian). 31: 380–393.
^ Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard, eds. (1 March 2015). "Genera in the family Gobiinae". Catalog of Fishes (genera). California Academy of Sciences. Retrieved 3 May 2015.
^ Neilson M.E.; Stepien C.A. (2009). "Escape from the Ponto-Caspian: Evolution and biogeography of an endemic goby species flock (Benthophilinae: Gobiidae: Teleostei)". Molecular Phylogenetics and Evolution. 52 (1): 84–102. Bibcode:2009MolPE..52...84N. doi:10.1016/j.ympev.2008.12.023. PMID 19475709.

External links

Neogobiin research at the University of Toledo, Ohio
Systematics of the Neogobiin Gobies Research project at U Toledo

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[1] Simonović P.D.; Nikolić V.P.; Skóra K.E. (1996). "Vertebral number in Ponto-Caspian gobies: phylogenetic relevance". J. Fish Biol. 49 (5): 1027–1029. Bibcode:1996JFBio..49.1027S. doi:10.1111/j.1095-8649.1996.tb00098.x.

[2] Miller P.J. (1986). "Gobiidae". In Whitehead P.J.P.; Bauchot M.-L.; Hureau J.-C.; Nielsen J.; Tortonese E. (eds.). Fishes of the North-eastern Atlantic and the Mediterranean Vol. 3. Paris: UNESCO.

[3] Pinchuk V.I. (1991). "K voprosu o grupirovkakh vidov v predelakh roda Neogobius (Perciformes)" [On the issue of species groupings within the genus Neogobius (Perciformes)]. Voprosy Ikhtiologii (in Russian). 31: 380–393.

[4] Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard, eds. (1 March 2015). "Genera in the family Gobiinae". Catalog of Fishes (genera). California Academy of Sciences. Retrieved 3 May 2015.

[5] Neilson M.E.; Stepien C.A. (2009). "Escape from the Ponto-Caspian: Evolution and biogeography of an endemic goby species flock (Benthophilinae: Gobiidae: Teleostei)". Molecular Phylogenetics and Evolution. 52 (1): 84–102. Bibcode:2009MolPE..52...84N. doi:10.1016/j.ympev.2008.12.023. PMID 19475709.

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